影片讲述松鼠奎特(克里斯·韦奇 Chris Wedge 配音)为了追松果,偶然引发了宇宙事件,改变并威胁着冰川时代的世界。为了拯救自己,话唠树懒希德(约翰·雷吉扎莫 John Leguizamo 配音)、猛犸象曼尼(雷·罗马诺 Ray Romano 配音)、剑齿虎迪亚哥(丹尼斯·利瑞 Denis Leary 配音),以及别的动物群族必须离开家园,踏上了他们充满喜剧色彩的冒险旅程,他们来到了充满异国情调的新大陆,并遇到了许多形形色色的新角色。
A hit man in black struggles into an apartment complex. It turns out that the apartments are hosts to a cornucopia of characters. A couple are about to separate or play baseball. A woman maintains a schedule of screaming in order for the neighbours to ascertain that she is alive. A mother just wants to have fun. A daughter just wants to obtain high marks and then someone walks on the roof looking for a UFO to act as a taxi.